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![PDF] Tribute to Tinbergen: The Four Problems of Biology. A Critical Appraisal | Semantic Scholar](https://d3i71xaburhd42.cloudfront.net/191c345b19e7b75b21f8683566e95ebc35fa4e98/2-Table1-1.png "PDF] Tribute to Tinbergen: The Four Problems of Biology. A Critical Appraisal | Semantic Scholar")
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PDF) Synthesis of Tinbergen's four questions and the future of sociogenomics
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![PDF] Tinbergen's four questions, organized: a response to Bateson and Laland. | Semantic Scholar](https://d3i71xaburhd42.cloudfront.net/6117ba207f35dbb9ef1c3c28d478e6b281e13d07/1-Table1-1.png "PDF] Tinbergen's four questions, organized: a response to Bateson and Laland. | Semantic Scholar")
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